A list of peer-reviewed papers from the Hedrick lab group and affiliates. This does not include preprints, conference proceedings; those can be found in Prof. Hedrick’s Google Scholar profile.


Zhang, C., Hedrick, T. L., & Mittal, R. (2018). An Integrated Study of the Aeromechanics of Hovering Flight in Perturbed Flows. AIAA Journal, 1-12.

Liang, Y., Urano, D., Liao, K. L., Hedrick, T. L., Gao, Y., & Jones, A. M. (2017). A nondestructive method to estimate the chlorophyll content of Arabidopsis seedlings. Plant methods, 13(1), 26.

Fernández, M. J., Driver, M. E., & Hedrick, T. L. (2017). Asymmetry costs: effects of wing damage on hovering flight performance in the hawkmoth Manduca sexta. J. Exp. Biol., 220(20), 3649-3656.

Brainerd, E. L., Blob, R. W., Hedrick, T. L., Creamer, A. T., & Müller, U. K. (2017). Data Management Rubric for Video Data in Organismal Biology. Integrative and comparative biology, 57(1), 33-47.

Evangelista, D. J., Ray, D. D., Raja, S. K., & Hedrick, T. L. (2017). Three-dimensional trajectories and network analyses of group behaviour within chimney swift flocks during approaches to the roost. Proc. R. Soc. B, 284(1849), 20162602.

Hedrick, T. L., Martínez-Blat, J., & Goodman, M. J. (2017). Flight motor modulation with speed in the hawkmoth Manduca sexta. Journal of insect physiology, 96, 115-121.

Jones, S. K., Yun, Y. J., Hedrick, T. L., Griffith, B. E., & Miller, L. A. (2016). Bristles reduce the force required to ‘fling’ wings apart in the smallest insects. J. Exp. Biol., 219(23), 3759-3772.

Cheng, B., Tobalske, B. W., Powers, D. R., Hedrick, T. L., Wang, Y., Wethington, S. M., … & Deng, X. (2016). Flight mechanics and control of escape manoeuvres in hummingbirds. II. Aerodynamic force production, flight control and performance limitations. J. Exp. Biol., 219(22), 3532-3543.

Cheng, B., Tobalske, B. W., Powers, D. R., Hedrick, T. L., Wethington, S. M., Chiu, G. T., & Deng, X. (2016). Flight mechanics and control of escape manoeuvres in hummingbirds. I. Flight kinematics. J. Exp. Biol., 219(22), 3518-3531.

Lohmann, A. C., Evangelista, D., Waldrop, L. D., Mah, C. L., & Hedrick, T. L. (2016). Covering ground: Movement patterns and random walk behavior in Aquilonastra anomala sea stars. The Biological Bulletin, 231(2), 130-141.

Warrick, D. R., Hedrick, T. L., Biewener, A. A., Crandell, K. E., & Tobalske, B. W. (2016). Foraging at the edge of the world: low-altitude, high-speed manoeuvering in barn swallows. Phil. Trans. R. Soc. B, 371(1704), 20150391.

Brace, N. L., Hedrick, T. L., Theriault, D. H., Fuller, N. W., Wu, Z., Betke, M., … & Morgansen, K. A. (2016). Using collision cones to assess biological deconfliction methods. Journal of The Royal Society Interface, 13(122), 20160502.

Jackson, B. E., Evangelista, D. J., Ray, D. D., & Hedrick, T. L. (2016). 3D for the people: multi-camera motion capture in the field with consumer-grade cameras and open source software. Biology open, 5(9), 1334-1342.

Song, J., Tobalske, B. W., Powers, D. R., Hedrick, T. L., & Luo, H. (2016). Three-dimensional simulation for fast forward flight of a calliope hummingbird. Royal Society open science, 3(6), 160230.

Greeter, J. S., & Hedrick, T. L. (2016). Direct lateral maneuvers in hawkmothsBiology Open, bio-012922.

Jones, S. K., Laurenza, R., Hedrick, T. L., Griffith, B. E., & Miller, L. A. (2015). Lift vs. drag based mechanisms for vertical force production in the smallest flying insects. Journal of theoretical biology384, 105-120.

Zhang, C., Hedrick, T. L., & Mittal, R. (2015). Centripetal acceleration reaction: An effective and robust mechanism for flapping flight in insectsPloS One10(8), e0132093.

Sholtis, K. M., Shelton, R. M., & Hedrick, T. L. (2015). Field flight dynamics of hummingbirds during territory encroachment and defensePLoS One10(6), e0125659.

Clifton, G. T., Hedrick, T. L., & Biewener, A. A. (2015). Western and Clark’s grebes use novel strategies for running on water. J. Exp. Biol.218(8), 1235-1243.

Song, J., Luo, H., & Hedrick, T. L. (2015). Wing-pitching mechanism of hovering ruby-throated hummingbirds. Bioinspiration & Biomimetics10(1), 016007.

Song, J., Luo, H., & Hedrick, T. L. (2015). Performance of a quasi-steady model for hovering hummingbirds. Theoretical and Applied Mechanics Letters5(1), 50-53.

Santhanakrishnan, A., Robinson, A. K., Jones, S., Low, A. A., Gadi, S., Hedrick, T. L., & Miller, L. A. (2014). Clap and fling mechanism with interacting porous wings in tiny insect flight. J. Exp. Biol.217(21), 3898-3909.

Hedrick, T. L., Combes, S. A., & Miller, L. A. (2014). Recent developments in the study of insect flightCanadian Journal of Zoology93(12), 925-943.

Song, J., Luo, H., Hedrick, T. L. (2014). Three-dimensional flow and lift characteristics of a hovering ruby-throated hummingbirdJ. R. Soc. Interface 11, 20140541 doi: 110.1098/​rsif.2014.0541.

Shelton, R. M., Jackson, B. E., Hedrick, T. L. (2014). The mechanics and behavior of cliff swallows during tandem flightsJ. Exp. Biol. doi: 10.1242/​jeb.101329.

Theriault, D. H., Fuller, N. W., Jackson, B. E., Bluhm, E., Evangelista, D., Wu, Z., Betke, M., Hedrick, T. L. (2014). A protocol and calibration method for accurate multi-camera field videographyJ. Exp. Biol. doi: 10.1242/​jeb.100529.

Ortega-Jimenez, V.M., Greeter, J.S.M., Mittal, R., Hedrick, T. L. (2013). Hawkmoth flight stability in turbulent vortex streets. J. Exp. Biol.doi: 10.1242/​jeb.089672.

Zheng, L., Hedrick, T. L., Mittal, R. (2013). A comparative study of the hovering efficiency of flapping and revolving wings. Bioinspiration & Biomimetics 8, 036001 doi:10.1088/1748-3182/8/3/036001.

Zheng, L., Hedrick, T. L., Mittal, R. (2013). A multi-fidelity modeling approach for evaluation and optimization of wing stroke aerodynamics in flapping flight. J. Fluid Mech. 721, 118-154 doi:10.1017/jfm.2013.46.

Zheng, L., Hedrick, T. L., Mittal, R. (2013). Time-Varying Wing-Twist Improves Aerodynamic Efficiency of Forward Flight in Butterflies. PLoS ONE 8(1), e53060 doi:10.1371/journal.pone.0053060.

Fernández, M. J., Springthorpe, D. and Hedrick, T. L. (2012). Neuromuscular and biomechanical compensation for wing asymmetry in insect hovering flight. J. Exp. Biol. 215, 3631-3638 doi: 10.1242/​jeb.073627.

Lin, T., Hedrick, T. L., Mittal, R. (2012). The significance of moment-of-inertia variation in flight manoeuvres of butterflies. Bioinspiration & Biomimetics 7, 044002 doi:10.1088/1748-3182/7/4/044002.

Springthorpe, D., Fernández, M. J. and Hedrick, T. L. (2012). Neuromuscular control of yaw turns in the hawkmoth J. Exp. Biol. 215, 1766-1774 doi: 10.1242/​jeb.067355.

Hedrick, T. L., Tobalske, B. W., Ros, I. G., Warrick, D. R., Biewener A. A. (2012). Morphological and kinematic basis of the hummingbird flight stroke: scaling of flight muscle transmission ratio. P. R. Soc. B. 279, 1986-1992 doi: 10.1098/rspb.2011.2238.

Hedrick, T. L. (2011). Damping in flapping flight and its implications for manoeuvring, scaling and evolution. J. Exp. Biol. 214, 4073-4081 doi: 10.1242/jeb.047001.

Cheng, B., Deng, X., Hedrick, T. L. (2011). The mechanics and control of pitching manoeuvres in a freely flying hawkmoth (Manduca sexta). J. Exp. Biol. 214, 4092-4106 doi: 10.1242/jeb.062760.

Hedrick, T. L., and Robinson, A.K. (2010). Within-wingbeat damping: dynamics of continuous free-flight yaw turns in Manduca sextaBiology Letters doi: 10.1098/rsbl.2010.0083.

Hedrick, T. L., Cheng, B. and Deng, X. (2009). Wingbeat time and the scaling of passive rotational damping in flapping flight. Science 5924, 252-255.

Hedrick, T. L. (2008). Software techniques for two- and three-dimensional kinematic measurements of biological and biomimetic systems. Bioinspiration & Biomimetics 3, 034001.